Inoculation experiments with Puccinia bromina Eriks., var. bromina urediospores collected from Bromus sterilis were unsuccessful on several grass species. However, aeciospores from Symphytum cordatum and S. tuberosum caused infections on Bromus erectus and B. ramosus subsp. benekenii. This supports the existence of two biological varieties: the non-alternating var. bromina and the heteroecious var. paucipora.
Urban Z., Marková-Ondráčková J. (1975): Inoculation experiments with Puccinia bromina Eriks. 2. – Česká Mykologie 29(3): 135–139.